Yes, Tim is right, the tape can certainly skew due to the path being out of whack, not just because a tape is edge curled or otherwise damaged. FWIW we don't see skewing too often in position 2 where the record head usually sits on a 1500. Position 4 can be a real booger sometimes. If it's skewing at pos 2 then the erase head in pos 1 might be misadjusted too.
Are the pinch rollers original or replacements? Some replacement rollers seem to cause some tracking issues. Look at the tape edge where it comes off the guide above the pinch roller, goes into the first roller, comes out of the second the roller, and goes over the guide above the second roller. Sometimes you can see a slight asymmetric curvature of the surface of the tape as it comes off those points and that may indicate problems around the headblock as well.